Arthropoda: Class Insecta

Class Insecta is the largest of the arthropod classes, containing hundreds of thousands of species. Except for a few primitive or highly modified forms, insects are characterized by having one or two pairs of wings attached to the thorax. The head region bears a pair of antennae, a pair of mandibles, and two pairs of modified maxillae forming the mouthparts. The abdomen is well set off from the thorax and has no appendages except reduced ones that are modified as reproductive organs. The typical insect head bears compound eyes and one or more simple eyes and is covered by a continuous exoskeletal armor. The thorax is made up of three segments, each bearing a pair of legs. The last two segments usually bear a pair of wings. Insects are predominantly terrestrial and have tracheae for gas exchange. Insects are also characterized by unique excretory organs, known as Malpighian tubules, which are useful in conserving water.

Members of the class are extremely varied. They have adapted to many different kinds of feeding and play a variety of important roles in their ecological communities. Mouthparts may be adapted to chewing either plant or animal food, for sucking plant sap or blood, or for lapping or swabbing moisture such as fruit juices or animal body fluids. Some burrow and feed in soil or plant tissue, some are runners or jumpers that feed at or near the ground level, and others feed on the wing.

Most primitive insects are wingless and have a relatively weak exoskeleton. These are forced to seek humid, protected habitats. Juveniles of primitive insects closely resemble the parents and undergo little change other than growth after hatching. This is called ametaboly. Many of the winged insects undergo paurometabolous development, hatching as nymphs that resemble the parent in many ways but that have small buds instead of wings. With each molt these juveniles change somewhat, and the wings increase in size as the young gradually assume the form of the adult. Some insects have adapted to an aquatic life to a certain extent, and in their juvenile stages they are found in ponds and streams. Some of these are hemimetabolous; the juveniles are naiads, i.e., they resemble the nymphs of paurometabolous insects, but their wings do not grow during the juvenile molts, even though other body changes occur. Instead, the last molt before the adult stage is reached involves full development of the wings, after which the insect takes up a terrestrial existence. The least primitive of the insects are termed holometabolous. In holometaboly, the eggs hatch to release the usually wormlike larvae, which are often equipped with false legs in the abdominal region to aid in locomotion. Wing buds are entirely lacking. Although the larvae grow at each molt, they do not begin to resemble the adult until later. During the larval stage the young insect enters into a quiescent pupal stage. At the end of this stage a major metamorphosis occurs, and the insect emerges with all the adult organs.

Insects often cause great losses in agriculture, attack stored products, parasitize humans and domesticated animals and plants, and serve as important carriers of disease organisms. They are also beneficial, producing honey and silk and pollinating the flowers of the majority of flowering plants.

Sections in this article:

• Introduction
• Class Insecta
• Class Symphyla
• Class Pauropoda
• Class Diplopoda
• Class Chilopoda
• Subphylum Crustacea
• Mandibulata
• Subphylum Chelicerata
• Subphylum Trilobitomorpha
• Bibliography

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